--------------o0o--------------

: Chính quy
Chuyên ngành
Khoa

: CNSH - CNTP
: 2011 - 2015


--------------o0o--------------

: Chính quy
Chuyên ngành
Khoa

: K43 - CNSH
: CNSH - CNTP
: 2011 - 2015


i

Công
S

C

,T

Nông Lâm Thái

................................................. 22
................... 36
................................... 38
......... 40

........................................................................................................ 42

................................................................................................. 43
...................... 45

................................................................ 47


iii

Trang
............................................................... 9
............................ 15
.............. 34
.................. 35
.................................................................... 39
....................... 40
................ 46


iv

CBD

: Cellulose binding domain

2

................................................................. 3
............... 3
......................................................................... 3
................................................................... 4
................................................................................. 8
.................................................................. 8

2.2.2. Enzyme cellulase..................................................................................... 9
........................ 15
..................................... 18
........................................................ 19
.......................................................... 19

......................................................................................................................... 20
........................................... 20
3.1.1.

............................................................ 20

3.1.2.

............................................................................... 20
......................................... 20
........................................................... 20


vi


vii

................................................................................. 47
5:

........................................................ 48
.................................................................................................... 48
.................................................................................................. 48


1

P

Ngày nay, ng

các

c

ngày

[26, 27].
nay

protease, amylase, pectinase, glucooxydase, cellulase.

cellula

vì

-

enzyme.
-

-

-

-

-

-

.


3

2.1.
2.1.1.

[12]

,

inh

,


m

Entodinium

nguyên sinh,
diplodinium

Caecomyces communis, Neocallimastix frontalis, Neocallimastix patriciarum,
Piromyces communis.
9

trong

và là tác nhân chính
ate


6

9

Trong

-1010

.
-

chính:

Fibrobacter succinogenes
Ruminococcus albus

[7].
[1]

3),
3

Tiêu hóa cellulose và hemicellulose

):
-D-

thành.
.


8

Hemicellulose
glucoronic, manose

enzyme cellulase
.

Cellulose => cellobiose
Cellobiose => glucose

[8].


enzyme
2.2.2

[29]

-

Aspergillus niger, Aspergillus oryzae

-

Actinomyces griseus, Streptomyces reticul

-

Acetobacter xylinum, Bacillus subtilis, Bacillus pumilis
- -Dlà glucose, cellobiose và cello-oligosaccharides.

2.2.2
: [20, 21, 22]
- Cellulobiohydrolase: enzyme
.K
-

- Endo- -D-

-49 kDa.
-1,4


2.2.3
Cellulase
hydroxyethyl cellulose (HEC).


12

-

-D-

-

-

500

-

0

-110 KDa
(Begunin,

.

2.2.2

a quá trình



Acetovibrio cellulolyticus

Lanzites trabeum

Clostridium cellulovorans

Poria placenta

Clostridium thermocellum

Tyromyces palustris

Phanerochaete chrysosporium

Butyrivibrio fibrisolvens

Sporotrichum thermonphile

Fibrobacter succinogenes

Coriolus versicolor

Ruminococcus albus

Caecomyces communis

Streptomyces lividans

Neocallimastix patriciarum

thành glucose.
Cellulose

C1

Cx

Cellulose

Cellobiose

-glucosidase

Glucose [8]

)

cellulase endoglucanase, e

-

thích khác nhau v
s và Recse (1964); Ogawa,
Toyam (1964); Iwasaki, Hayoshi, Walabayais, Nisizwa (1964); Funata
(1965); King (1965) và Srinivasan (1973) [6].


15

[6].

-

11, 13].